Parks and Recreation fans are still reeling that human plague Jerry Gergich was revealed to have a lovely, stone-cold fox of a daughter in Millicent. Now the office doormat will finally introduce us to his wife, Gayle, as played by Christie Brinkley, former Playboy model, Sports Illustrated Swimsuit Issue cover girl, and onetime wife of Billy Joel. She’ll appear in the fifth episode, one that will feature more Lucy Lawless and the return of Megan Mullally.